After a family tragedy turns her life upside down, 16-year-old high schooler Tooru Honda takes matters into her own hands and moves out...into a tent! Unfortunately for her, she pitches her new home on private land belonging to the mysterious Souma clan, and it isn't long before the owners discover her secret. But, as Tooru quickly finds out when the family offers to take her in, the Soumas have a secret of their own--when hugged by the opposite sex, they turn into the animals of the Chinese Zodiac!

High schooler Yuri finds herself atop a skyscraper in a strange world, where she must survive against masked assailants bent on killing their prey.

Long ago, a war broke out in Japan that pitted Kantou against Kansai and divided the country. Kansai became dependent on Kantou, but slowly the police and government began losing control of Kansai to criminals known as Akudama.

The story is set in the distant future. The land is ruined, and humanity establishes the mobile fort city Plantation. Pilots produced inside Plantation live in Mistilteinn, also know as the "birdcage." Children live there knowing nothing of the outside world or the freedom of the sky. Their lives consist of battling to carry out missions. Their enemies are mysterious giant lifeforms known as Kyouryuu, and the children pilot robots called Franxx to face off against them. For the children, riding the Franxx proves their existence. A boy named Hiro is called Code:016, and he was once known as a prodigy. However, he has fallen behind, and his existence seems unnecessary. Not piloting a Franxx is the same as ceasing to exist. One day, a mysterious girl known as "Zero Two" appears before him. Two horns grow out of her head.